Usually means: Having openings or perforations present.
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We found 4 dictionaries that define the word aperturate:

General (2 matching dictionaries)
  1. aperturate: Wordnik
  2. aperturate: Wiktionary

Science (2 matching dictionaries)
  1. Archaeology Wordsmith (No longer online)
  2. Bryological (No longer online)

Definitions from Wiktionary (aperturate)

adjective:  (palynology, of a pollen grain) Having one or more apertures

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