Usually means: Spoke in a low, angry tone.
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We found 14 dictionaries that define the word growled:

General (12 matching dictionaries)
  1. growled: Merriam-Webster
  2. growled: Collins English Dictionary
  3. growled: Vocabulary.com
  4. growled: Wordnik
  5. growled: Cambridge Advanced Learner's Dictionary
  6. growled: Wiktionary
  7. Growled, growled: Dictionary.com
  8. growled: Cambridge Essential American English Dictionary
  9. Growled: Online Plain Text English Dictionary
  10. Growled: AllWords.com Multi-Lingual Dictionary
  11. growled: FreeDictionary.org
  12. growled: TheFreeDictionary.com

Computing (1 matching dictionary)
  1. growled: Encyclopedia

Miscellaneous (1 matching dictionary)
  1. growled: Idioms

(Note: See growl as well.)

Definitions from Wiktionary (growl)

noun:  A deep, rumbling, threatening sound made in the throat by an animal.
noun:  (by extension) A similar sound made by a human.
noun:  (by extension) The rumbling sound made by a human's hungry stomach.
noun:  (by extension) An aggressive grumbling.
noun:  (jazz, by extension) A low-pitched rumbling sound produced with a wind instrument.
noun:  (by extension) Death growl
verb:  (intransitive) To utter a deep guttural sound, as an angry animal; to give forth an angry, grumbling sound.
verb:  (intransitive, jazz) Of a wind instrument: to produce a low-pitched rumbling sound.
verb:  (intransitive, software) To send a user a message via the Growl software library.
verb:  (transitive) To express (something) by growling.
verb:  (transitive, jazz) To play a wind instrument in a way that produces a low-pitched rumbling sound.
verb:  (intransitive) To perform death growl vocals.
▸ Also see growl


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